Publication Date

7-9-2024

Journal

Proceedings of the National Academy of Sciences of the United States of America

DOI

10.1073/pnas.2404062121

PMID

38968109

PMCID

PMC11253012

PubMedCentral® Posted Date

7-5-2024

PubMedCentral® Full Text Version

Post-print

Published Open-Access

yes

Keywords

Trophoblasts, Humans, Basic Helix-Loop-Helix Leucine Zipper Transcription Factors, Female, Pregnancy, Mice, Animals, Cell Differentiation, Mechanistic Target of Rapamycin Complex 1, Placenta, Signal Transduction, Autophagy, syncytiotrophoblast, TFEB, ERVFRD-1, human trophoblast stem cell, fetal growth restriction

Abstract

The formation of multinucleated syncytiotrophoblast (STB) through cell fusion of cytotrophoblast, also termed syncytialization, ensures the proper placental structure and functions. Nutrient insufficiency and inactivation of the mechanistic target of rapamycin complex 1 (mTORC1) in trophoblasts have been shown to enhance STB formation; however, the underlying mechanism remains elusive. Here, we showed that the deficiency of a mTORC1 downstream transcriptional factor, TFEB, significantly impaired STB formation in human trophoblasts and knock-out mice. TFEB conferred direct transcriptional activation of the fusogen ERVFRD-1 and thereby promoted trophoblast syncytialization. Additionally, TFEB expression positively correlated with the reinforced trophoblast syncytialization in human fetal growth restriction placentas with suppressed mTORC1 activity. Our findings substantiate that the TFEB-fusogen axis safeguards proper STB formation during placenta development.

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