Publication Date

7-10-2024

Journal

Cell Genomics

DOI

10.1016/j.xgen.2024.100590

PMID

38908378

PMCID

PMC11293582

PubMedCentral® Posted Date

6-21-2024

PubMedCentral® Full Text Version

Post-print

Published Open-Access

yes

Keywords

Humans, Haplotypes, Comparative Genomic Hybridization, Genomic Structural Variation, Genome, Human, Gene Duplication, copy-number variant, break-induced replication, recombination, Xq28, inversions, segmental duplication, MECP2 duplication syndrome, Mendelian diseases, MMBIR, template switching

Abstract

The duplication-triplication/inverted-duplication (DUP-TRP/INV-DUP) structure is a complex genomic rearrangement (CGR). Although it has been identified as an important pathogenic DNA mutation signature in genomic disorders and cancer genomes, its architecture remains unresolved. Here, we studied the genomic architecture of DUP-TRP/INV-DUP by investigating the DNA of 24 patients identified by array comparative genomic hybridization (aCGH) on whom we found evidence for the existence of 4 out of 4 predicted structural variant (SV) haplotypes. Using a combination of short-read genome sequencing (GS), long-read GS, optical genome mapping, and single-cell DNA template strand sequencing (strand-seq), the haplotype structure was resolved in 18 samples. The point of template switching in 4 samples was shown to be a segment of ∼2.2-5.5 kb of 100% nucleotide similarity within inverted repeat pairs. These data provide experimental evidence that inverted low-copy repeats act as recombinant substrates. This type of CGR can result in multiple conformers generating diverse SV haplotypes in susceptible dosage-sensitive loci.

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