Publication Date

4-24-2023

Journal

Nature Communications

DOI

10.1038/s41467-023-37984-x

PMID

37095084

PMCID

PMC10125973

PubMedCentral® Posted Date

4-24-2023

PubMedCentral® Full Text Version

Post-print

Published Open-Access

yes

Keywords

Male, Mice, Animals, Sperm Maturation, Semen, Testis, Epididymis, Spermatozoa, Fertility, Mammals, Spermatogenesis, Extracellular signalling molecules, Reproductive biology

Abstract

The mammalian spermatozoa produced in the testis require functional maturation in the epididymis for their full competence. Epididymal sperm maturation is regulated by lumicrine signalling pathways in which testis-derived secreted signals relocate to the epididymis lumen and promote functional differentiation. However, the detailed mechanisms of lumicrine regulation are unclear. Herein, we demonstrate that a small secreted protein, NELL2-interacting cofactor for lumicrine signalling (NICOL), plays a crucial role in lumicrine signalling in mice. NICOL is expressed in male reproductive organs, including the testis, and forms a complex with the testis-secreted protein NELL2, which is transported transluminally from the testis to the epididymis. Males lacking Nicol are sterile due to impaired NELL2-mediated lumicrine signalling, leading to defective epididymal differentiation and deficient sperm maturation but can be restored by NICOL expression in testicular germ cells. Our results demonstrate how lumicrine signalling regulates epididymal function for successful sperm maturation and male fertility.

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