Publication Date

11-1-2023

Journal

Journal of Medical Genetics

DOI

10.1136/jmg-2022-109127

PMID

37316189

PMCID

PMC11206234

PubMedCentral® Posted Date

11-1-2024

PubMedCentral® Full Text Version

Author MSS

Published Open-Access

yes

Keywords

Developmental Disabilities, Ikaros Transcription Factor, Hearing Loss, Syndrome, Humans, DNA-Binding Proteins, Craniofacial Abnormalities, Breast Diseases, l IKZF2, Helios, bilateral sensorineural hearing loss, immune dysregulation, craniofacial differences, syndrome, exome sequencing, genome sequencing

Abstract

Background:

Helios (encoded by IKZF2), a member of the Ikaros family of transcription factors, is a zinc finger protein involved in embryogenesis and immune function. Although predominantly recognized for its role in the development and function of T lymphocytes, particularly the CD4+ regulatory T cells (Tregs), the expression and function of Helios extends beyond the immune system. During embryogenesis, Helios is expressed in a wide range of tissues, making genetic variants that disrupt the function of Helios strong candidates for causing widespread immune-related and developmental abnormalities in humans.

Methods:

We performed detailed phenotypic, genomic, and functional investigations on two unrelated individuals with a phenotype of immune dysregulation combined with syndromic features including craniofacial differences, sensorineural hearing loss, and congenital abnormalities.

Results:

Genome sequencing revealed de novo heterozygous variants that alter the critical DNA-binding zinc fingers (ZF) of Helios. Proband 1 had a tandem duplication of ZFs 2 and 3 in the DNA-binding domain of Helios (p.Gly136_Ser191dup) and Proband 2 had a missense variant impacting one of the key residues for specific base recognition and DNA interaction in ZF2 of Helios (p.Gly153Arg). Functional studies confirmed that both these variant proteins are expressed, and that they interfere with the ability of the wild-type Helios protein to perform its canonical function—repressing IL2 transcription activity—in a dominant negative manner.

Conclusion:

This study is the first to describe dominant negative IKZF2 variants. These variants cause a novel genetic syndrome characterized by Immunodysregulation, Craniofacial anomalies, Hearing impairment, Athelia, and Developmental delay (ICHAD syndrome).

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