Language

English

Publication Date

10-1-2025

Journal

EMBO Molecular Medicine

DOI

10.1038/s44321-025-00290-8

PMID

40859057

PMCID

PMC12514044

PubMedCentral® Posted Date

8-26-2025

PubMedCentral® Full Text Version

Post-print

Abstract

ATP5F1A encodes the α-subunit of complex V of the respiratory chain, which is responsible for mitochondrial ATP synthesis. We describe 6 probands with heterozygous de novo missense ATP5F1A variants that presented with developmental delay, intellectual disability, and movement disorders. All variants were located at the contact points between the α- and β-subunits. Functional studies in C. elegans revealed that the variants were damaging via a dominant negative genetic mechanism. Biochemical and proteomics studies of proband-derived cells showed a marked reduction in complex V abundance and activity. Mitochondrial physiology studies revealed increased oxygen consumption, yet decreased mitochondrial membrane potential and ATP levels indicative of uncoupled oxidative phosphorylation as a pathophysiologic mechanism. Our findings contrast with the previously reported ATP5F1A variant, p.Arg207His, indicating a different pathological mechanism. This study expands the phenotypic and genotypic spectrum of ATP5F1A-associated conditions and highlights how functional studies can provide an understanding of the genetic, molecular, and cellular mechanisms of ATP5F1A variants of uncertain significance. With 12 heterozygous individuals now reported, ATP5F1A is the most frequent nuclear genome cause of complex V deficiency.

Keywords

Humans, Oxidative Phosphorylation, Male, Female, Animals, Mitochondrial Proton-Translocating ATPases, Caenorhabditis elegans, Mutation, Missense, Nervous System Diseases, Child, Mitochondria, Child, Preschool, ATP5F1A, ATP Synthase, Complex V, Mitochondriopathy, Oxidative Phosphorylation, Genetics, Gene Therapy & Genetic Disease; Neuroscience; Organelles

Published Open-Access

yes

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