Language

English

Publication Date

7-1-2026

Journal

Genes & Development

DOI

10.1101/gad.353584.125

PMID

42156139

PMCID

PMC13267984

PubMedCentral® Posted Date

7-1-2026

PubMedCentral® Full Text Version

Post-print

Abstract

Differentiation requires coordinated exit from the stem cell state, during which gene regulatory networks sustaining self-renewal are dismantled, while lineage-specific programs are activated. This transition is governed by chromatin modifications, transcriptional networks, RNA processing, translational control, and metabolic rewiring that must operate with temporal precision. Despite significant progress in identifying individual regulatory components, understanding how these layers integrate to orchestrate irreversible cell fate commitment remains a fundamental challenge. This review examines common and unique regulatory principles governing stem cell exit, from totipotency during early embryogenesis to tissue-specific stem cell differentiation in adults. We synthesize recent findings on regulatory mechanisms across mammalian species, highlight species-specific adaptations, and explore the concept of reversibility in differentiation. Elucidating these principles has broad implications for regenerative medicine, cellular reprogramming, and diseases in which differentiation programs are corrupted.

Keywords

Animals, Cell Differentiation, Stem Cells, Humans, Mammals, Gene Expression Regulation, Developmental, Gene Regulatory Networks, Cell Lineage, Chromatin, cell fate, chromatin, epigenetics, pluripotency, RNA processing, stem cells

Published Open-Access

yes

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