Language

English

Publication Date

2-1-2026

Journal

Nature Neuroscience

DOI

10.1038/s41593-025-02166-z

PMID

41436651

PMCID

PMC13032191

PubMedCentral® Posted Date

6-30-2026

PubMedCentral® Full Text Version

Author MSS

Abstract

Parcellation of the cerebral cortex into functionally modular brain areas is foundational to cognitive and systems neuroscience. Here, we question the central status of brain areas from the perspectives of neuroanatomy and electrophysiology. We argue that the major ostensible determinants of brain function, such as cytoarchitecture and connectivity, seldom produce convergent parcellations. Brain areas themselves are just one of several equally important organizing principles; others include macroscale gradients, distributed networks, layers, columns and patches. We further argue that the evidence for a close correspondence between areal parcellation and cognitive function is weaker than is generally supposed. Indeed, many important cognitive functions appear to be implemented in a broadly distributed manner, whereas others appear to obey organizations that have little relationship to brain areas, including distributed networks and functional gradients. We conclude by suggesting a set of guiding principles for performing systems and cognitive neuroscience without the intellectual foundation provided by arealization.

Keywords

Humans, Brain, Animals, Cognition, Brain Mapping, Nerve Net, Neural Pathways, Cerebral Cortex, neuroanatomy, arealization, localization, parcellation, cerebral cortex, mass action

Published Open-Access

yes

Share

COinS
 
 

To view the content in your browser, please download Adobe Reader or, alternately,
you may Download the file to your hard drive.

NOTE: The latest versions of Adobe Reader do not support viewing PDF files within Firefox on Mac OS and if you are using a modern (Intel) Mac, there is no official plugin for viewing PDF files within the browser window.