Author ORCID Identifier

0009-0002-8656-3869

Date of Graduation

8-2024

Document Type

Thesis (MS)

Program Affiliation

Genetics and Epigenetics

Degree Name

Masters of Science (MS)

Advisor/Committee Chair

Margarida Santos, PhD

Committee Member

Mark T. Bedford, PhD

Committee Member

Xiaodong Cheng, PhD

Committee Member

Joya Chandra, PhD

Committee Member

Deepa Sampath, PhD

Committee Member

M. James You, MD, PhD

Abstract

Myeloproliferative neoplasms (MPNs) are a hematopoietic disease characterized by hyperproliferation of cells of the myeloid lineage for which current therapeutic options are limited. Discovered in 2005, the JAK2V617F mutation is the most common driver mutation in BCR-ABL negative MPNs, resulting in constitutive activation of the JAK2 protein and the JAK-STAT signaling pathway. A role for the methyltransferase activity of Protein Arginine Methyltransferase 5 (PRMT5) has been proposed in JAK2-mutant MPN, highlighting both a mechanism through which this mutation can drive disease progression and a potential mode of therapeutic intervention. Staphylococcal Nuclease Domain-Containing Protein 1 (SND1) is the effector molecule responsible for reading the symmetric dimethyl-arginine (SDMA) marks deposited by PRMT5, underpinning a potential critical role in MPN pathogenesis and treatment. SND1 is the only human protein known to contain the Staphylococcal nuclease-like domain, making it a uniquely promising therapeutic target for which there is little potential for off-site effects by small molecule inhibition. Using a human JAK2V617F-mutant MPN cell line as a disease model, we are actively exploring the roles of PRMT5 and SND1 in the progression of this class of diseases. We are establishing the effects that reduced or abolished PRMT5/SND1 activity have on cell fitness by generating genetically modified cell lines and utilizing small molecule inhibitors. Both SND1 and PRMT5 show promise in altering the proliferation and differentiation patterns of JAK2-mutant MPN cells, encouraging further investigation into their potential as therapeutic targets in the treatment of these diseases.

Keywords

PRMT5, SND1, JAK2, myeloproliferative neoplasms

Available for download on Friday, August 08, 2025

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