Author ORCID Identifier

https://orcid.org/0000-0001-5430-5799

Date of Graduation

12-2024

Document Type

Dissertation (PhD)

Program Affiliation

Genetics and Epigenetics

Degree Name

Doctor of Philosophy (PhD)

Advisor/Committee Chair

Dr. Richard Behringer

Committee Member

Dr. Vicki Huff

Committee Member

Dr. George Eisenhoffer

Committee Member

Dr. Rachel Miller

Committee Member

Dr. Yoshihiro Komatsu

Abstract

WT1 is a zinc finger transcription factor widely expressed in the urogenital system. Human mutations of WT1 lead to pediatric nephroblastoma as well as frequent differences of sex development (DSDs). Previous studies have suggested that WT1 acts as an activator for Amhr2, a necessary component of typical male differentiation. We used the mouse as a model to investigate the role of WT1 in sex development, where we deleted Wt1 in the Müllerian duct (MD) mesenchyme using a novel conditional null reporter allele, Wt1 flox-RFP. This allele utilizes the Cre-lox system to delete exons 8 and 9 of Wt1, disrupting the DNA binding capability of the protein. Simultaneously, a nuclear red fluorescent protein is activated in cells expressing the recombined allele. After validating and characterizing this allele, we drove Wt1 deletion in the MD mesenchyme using Amhr2-Cre. We found that loss of WT1 function in the MD mesenchyme resulted in the male developing a uterus. This is the first in vivo demonstration that Wt1 has a role in MD regression within the MD mesenchyme.

Keywords

Wt1, development, genetics, uterus, oviduct, DSD, Amhr2, Mullerian duct

Available for download on Saturday, August 30, 2025

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