Author ORCID Identifier

https://orcid.org/0000-0003-2722-0081

Date of Graduation

12-2025

Document Type

Dissertation (PhD)

Program Affiliation

Immunology

Degree Name

Doctor of Philosophy (PhD)

Advisor/Committee Chair

Balveen Kaur

Committee Member

Louise McCullough

Committee Member

Michael Curran

Committee Member

Kapil Bhalla

Committee Member

Jason Huse

Abstract

Oncolytic HSV (oHSV) therapy is a live virus-based immunotherapy that lyses tumor cells which release antigens and activate anti-tumor immunity. Immunotherapy-based drugs have made significant advances in treating cancer. However, unchecked immune activation can escalate to autoreactivity or toxicity, so this inflammation is frequently accompanied by anti-inflammatory responses to maintain tissue homeostasis. Tumor cells can exploit anti-inflammatory pathways to escape anti-tumor immune responses. Here we identify that oHSV immune therapy rewires the tumor microenvironment to take advantage of immunosuppressive extracellular adenosine (eADO) signaling as a means of escaping host anti-tumor immune responses. Transcriptomic analysis of patient tumors pre- and post-virotherapy with CAN-3110 revealed increased expression of the adenosine receptor gene ADORA2B after treatment. The accumulation of eADO in the TME is predominantly the responsibility of ectoenzyme CD73 (NT5E) which catalyzes the hydrolysis of AMP to eADO. High NT5E gene expression, as well as gene signatures suggestive of adenosine signaling, correlated with a significantly worse prognosis for patients with solid tumors. Here, we examine the impact of blocking CD73 on therapeutic response to oHSV in immune competent solid tumor models. Tumor-bearing CD73 global knock-out mice treated with oHSV and wild type mice treated with a murine CD73 antibody and oHSV showed significant improvements in therapeutic efficacy of oHSV. Single cell RNA sequencing was employed to assess changes in immune cell infiltration and communication. Flow cytometric immunophenotyping and immunofluorescent imaging were utilized to confirm single cell sequencing predicted changes in tumor microenvironment. Sequencing data and in vivo blockade of myeloid cell recruitment into tumors revealed that this beneficial therapeutic effect partially relies on increased macrophage mediated antigen presentation. The combination of oHSV with a function-blocking CD73 antibody supports the development of an anti-tumor immune response in solid tumors. We then isolated and characterized a novel, function blocking, human CD73 antibody by screening a human phage display library. This antibody reduces virus-induced eADO and improves anti-tumor responses of cultured human immune cells. To evaluate the efficacy of Ab6, we created humanized mice and observed significant anti-tumor efficacy of Ab6 in combination with oHSV. These studies support the further development of Ab6 for safety and efficacy studies in patients.

Keywords

oncolytic virus therapy, oHSV therapy, oHSV, oncolytic herpes virus, CD73, adenosine signaling, solid tumor, glioblastoma, tumor microenvironment, antibody, immune therapy

Available for download on Saturday, December 12, 2026

Share

COinS